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, (. Pantopoda, Gerstaeker, 1862) (Chelicerata)[1]. , , [2]. . 1000 [3]. [4].
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- Pycnogonida Latreille, 1810
- Podosomata Lh, 1815
- Phalangium Lineus, 1767[5].
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() . (Nympnon sp.) (Pycnogonium sp.) . - . 1-72 ; 1,4 50 .
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( 5-6) , . «» . , Y- , ( ), , . , , . , . , (. ).
. , , -, . . , . (, Anoplodactylus petiolatus) , . . , , (Nymphon procerus 2500 6000). Nymphon longicoxa, Heteronymphon sp., , . , , , , , , .
, , . 2-3 . (Nymphon sp.), , . , (Achelia sp.). (Pycnogonum sp.). , , (Pseudopallene sp.). , .
, , . . , (Achelia sp.) (Neopallene sp.) , (Phoxichilidium sp., Pallene sp.). , . . .
, , , . . , . Phoxichilidium sp. , Nymphon sp. , , . , . . .
, . . Pantopoda (Pentanymphon sp., Pentapycnon sp., Decalopoda sp.) (Dodecalopoda sp) .
. «coxae» , ; «femur»; «tibia» ; «tarsus» «propodus». . . , . , (), ().
. . . , .
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. . . . .
, , Palaeisopus problematicus Palaeopantopus maucheri, , .
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Y- , . , . . . Pseudopallene spinipes . , . c , , . . , .
. . . . , - B, - . . , , . .
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, , (), ().
, (), , . , . (Pseudopallene spinipes) (Nymphon rubrum) () . , . . . , , , , .
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, . , . , ( ). M. Heb, R.R.Melzer U.Smola Anoplodacylus petiolatus (. Phoxichilidiidae). ( «»). . , . , . . ( ). , . , - . , .
, , . «-», . Nymphon sp. Pycnogonum sp. , , ( Pycnogonum litorale 15 lux).
.
, 23 , . 2- 3- 4- 5- .
; , ; 47- , 2- , 6- 7- ( 5 ) ; , 4- ; 47- . , , , , .
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. ( ). , . ( (Polychaeta), (Foraminifera), (Bryozoa), (Ciliophora), (Porifera) .). , () . , .
, , (propodus) . , , . , «», - (coxa1 coxa2) .
. (Polychaeta), (Bryozoa), (Cnidaria), (Nudibranchia), (Crustacea), (Holothuroidea). , . , . , . (Amothella sp., Fragilia sp., Heterofragilia sp), (Eurycyde sp., Ephyrogymma sp.) (Tanystilla sp.) . ( ). .
Nymphon, Pseudopallene , , ( ) . .
. , , . , Anoplodactylus ophiurophilus Ophiocoma , , . , Ophiocoma, , , .
(Lecythorhynchus hilgendorfi Holothuria lubrica ). . Nymphonella tapetis Paphia philippinarum. , 7-10 , . , .
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Pantopoda . .
. 1802 , Pycnogonides Arachnida, , . , , .
, 1890 , Pantopoda , (, , , ). , .
, 1899 , , . , . , , . , , , Chelicerata . , ( ). . , Chelicerata , . . . , , . . . . , (Pentanymphon sp.) (Dodecalopoda sp.) , . , , - . , . . Chelicerata Euchelicerata (Merostomata Chelicerata s.s.) Pantopoda. ,
, . EF-1a, EF-2, II, 18S 28S , 18S D3 28S , H3 U2 .
Crustacea. XIX . , , , . Araneiformes. () . , Crustacea Pantopoda , . . , (), , (). , . ( ), ( ). , , , . , .
. XX Zrzavy (1998). , Pantopoda , Chelicerata Pycnogonida . 10 .
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6
- Cambropycnogonon klausmulleri Waloszek, Dunlop, 2002 .
- Haliestes dasos Siveter et al., 2004 .
- Palaeisopus problematicus Broili, 1928 .
- Palaeopantopus maucheri Broili, 1929 .
- Palaeothea devonica Bergstrom et al., 1980 .
- Flagellopantopus blocki Poschmann, Dunlop, 2006 .
(Dunlop & Arango, 2005) Pycnogonida Arachnomorpha ( + ) Euarthropoda Chelicerata. . 1300 68 , :
- Ammotheidae
- Austrodecidae
- Callipallenidae
- Colossendeidae
- Endeididae
- Endeididae
- Nymphonidae
- Pallenopsidae
- Phoxichilidiidae
- Pycnogonidae
- Rhynchothoracidae
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- Pycnogonum , , , ; Pantopoda littorale ;
- Nymphon , ; Pantopoda longitarse . ;
- Collossendeis , . Pantopoda proboscidea 8 , 30 , . .
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- Dunlop J. A., Arango C. P. Pycnogonid affinities: a review // Blackwell Verlag, Berlin, 2004,P. 8-21.
- Arnaund F. and R. N. Bamber, 1987. The biology of Pycnogonida // Advances in Marine Biology. Vol. 24. P. 1 96.
- .., (Pantopoda), , 1929, . 1-555.
- Zrzavy J. Phylogeny of the Metazoa Based on Morphological and 18S Ribosomal DNA Evidence // Cladistics 14, 1998
- Morgan T. H., 1891. A contribution to the embryology and phylogeny of the pycnogonids // Studies from the Biological Laboratory of Johns Hopkins University. Baltimore. Vol. 5. No. 1. P. 1 76.
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- Cotton, T.J., & Braddy, S.J. (2004). The phylogeny of arachnomorph arthropods and the origin of the Chelicerata. // Transactions of the Royal Society of Edinburgh: Earth Sciences, 94: 169193. DOI:10.1017/S0263593300000596 (.)
- Dunlop J. A. & C. P. Arango (2005). Pycnogonid affinities: a review. // Journal of Zoological Systematics and Evolutionary Research 43: 8-21. DOI:10.1111/j.1439-0469.2004.00284.x (.)
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|---|---|
| · Eumetazoa · Bilateria · Protostomia · Ecdysozoa | |
| (Chelicerata) | (Arachnida) · (Merostomata) · (Pantopoda) |
| (Myriapoda) | (Chilopoda) · (Diplopoda) · (Pauropoda) · (Symphyla) |
| (Hexapoda) | (Insecta) · (Entognatha) |
| (Crustacea) | (Branchiopoda) · (Remipedia) · (Cephalocarida) · (Maxillopoda) · (Ostracoda) · (Malacostraca) |